| Holder | Shares | % out | Change |
|---|---|---|---|
| Merck & Co., Inc. | 1,214,126 | 14.6% | — |
| Ikarian Capital, LLC | 74,698 | 0.9% | +0 |
| Rathbones Group PLC | 45,723 | 0.5% | +0 |
| CITADEL ADVISORS LLC | 36,584 | 0.4% | +18,811 |
| RENAISSANCE TECHNOLOGIES LLC | 33,800 | 0.4% | −5,425 |
| Annis Gardner Whiting Capital Advisors, LLC | 30,000 | 0.4% | +0 |
| J. Safra Sarasin Holding AG | 22,896 | 0.3% | — |
| RHUMBLINE ADVISERS | 21,849 | 0.3% | +190 |
| JANE STREET GROUP, LLC | 20,581 | 0.2% | — |
| BOOTHBAY FUND MANAGEMENT, LLC | 18,426 | 0.2% | +0 |
| Stonepine Capital Management, LLC | 17,562 | 0.2% | −2,151 |
| SmartHarvest Portfolios, LLC | 13,460 | 0.2% | −348 |
| XTX Topco Ltd | 12,899 | 0.2% | −9,154 |
| WESBANCO BANK INC | 10,000 | 0.1% | +0 |
| NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO | 8,000 | 0.1% | +0 |